Skip to main content


Information on this website is posted for historical reference only. Please visit the Office of the Registrar for current requirements.


Chair: Hermes H. Yeh

Professors D. Bartlett Jr., R. A. Darnall (Pediatrics and Physiology), J. A. Daubenspeck (Physiology and Biomedical Engineering), G. Fejes-Toth, V. A. Galton, P. M. Guyre, L. P. Henderson (Physiology and Biochemistry), J. C. Leiter (Physiology and Medicine), H. L. Manning (Medicine and Physiology), R. A. Maue (Physiology and Biochemistry), A. Naray-Fejes-Toth, E. E. Nattie, W. G. North, D. L. St. Germain (Medicine and Physiology), B. A. Stanton, H. M. Swartz (Radiology, Physiology and Community and Family Medicine), C. R. Wira, H. H. Yeh; Visiting Professor A. Katz (Medicine and Physiology); Associate Professors E. deMuinck (Medicine and Physiology), R. B. Robey; Assistant Professor A. Gulledge; Research Associate Professors J. E. Bodwell, A. L. Givan; Research Assistant Professors J.V. Fahey, A. Li, P. A. Pioli, L. A. Sheldon.

The Molecular, Cellular and Systems* Physiology Graduate Program is centered in the Physiology Department of Dartmouth Medical School, which is located in the Borwell Building at the Dartmouth-Hitchcock Medical Center. It includes faculty from other departments in the Medical School. The courses listed below are designed for graduate students.

*As of the 2007-2008 academic year the MCSP program has merged with the new Program in Experimental and Molecular Medicine, and thus the courses listed pertain only to students in year 2 and beyond.


To qualify for award of the Ph.D. degree, a student must fulfill the following requirements:

1. The Medical Physiology courses (Physiology 110 and 120) plus an additional section in Gastro-Intestinal Physiology.

2. Two terms of Biochemistry.

3. The Medical Neuroscience course.

4. The Critical Reading of Scientific Literature course.

5. A course on Scientific Ethics.

6. Three research rotations.

7. Two Advanced Physiology courses.

8. Two elective courses, which may be either a core course or an advanced course offered by either Physiology or another department.

9. Thesis research propositional examination.

10. Laboratory research leading to the preparation of a thesis.

11. Defense of the thesis.