Darren M. Ward Department of Biological Sciences
Dartmouth College

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darren dot ward at dartmouth dot edu


I do research in basic and applied aquatic ecology.  My primary research interests are population dynamics, predator/prey interactions, and trace element cycling in food webs.  Most of my research has a strong field work component because that's the fun part of ecology!  

Recent and ongoing projects:

  • Mercury accumulation in stream food webs
  • Atlantic salmon restoration
  • Eurasian watermilfoil biological control

Mercury in streams

Mercury frequency distributionMercury accumulation in fish is an ongoing problem.  We developed a unique approach to measure standardized mercury accumulation rates for free-living fish in New England streams.  In spring, we stock juvenile fish from a uniform source population in multiple streams.  The fish then grow and accumulate mercury over the growing season.  In the fall, we collect samples to measure mercury concentrations.  We have applied this approach to identify important ecological determinants of mercury accumulation in stream fish.

Relevant publications:

Ward, D.M., K.H. Nislow, C.Y. Chen, and C.L. Folt.  2010.  Rapid, efficient growth reduces mercury concentrations in stream-dwelling Atlantic salmon.  Transactions of the American Fisheries Society 139:1-10.

Atlantic salmon restoration

Atlantic salmon parr, juvenile Atlantic salmonAtlantic salmon were eliminated from the Connecticut River basin in New England about 200 years ago when dams blocked their spawning migrations. Now, we are trying to reestablish an Atlantic salmon population in the Connecticut River.  My dissertation research addressed factors that limit survival and growth of reintroduced juvenile Atlantic salmon in Connecticut River tributaries.


Slimy sculpin Some of my dissertation work examined the effects of slimy sculpin on juvenile Atlantic salmon survival.  These small, inconspicuous fish turned out to be important predators of stocked salmon fry.

Relevant publications:

Ward, D.M., K.H. Nislow, C.L. Folt.  2009.  Increased population density and suppressed prey biomass: relative impacts on juvenile Atlantic salmon growth.  Transactions of the American Fisheries Society 138:135-143.

Ward, D.M., K.H. Nislow, and C.L. Folt.  2008.  Predators reverse the direction of density dependence for juvenile salmon mortality. Oecologia 156:515-522.

Ward, D.M., K.H. Nislow, and C.L. Folt. 2008.  Do native species limit survival of reintroduced Atlantic salmon in historic rearing streams? Biological Conservation 141:146-152.

Ward, D.M., K.H. Nislow, J.D. Armstrong, S. Einum, and C.L. Folt. 2007. Is the shape of the density-growth relationship for stream salmonids evidence for exploitative rather than interference competition? Journal of Animal Ecology 76: 135-138.

 Eurasian watermilfoil control- research page at the University of Minnesota

Milfoil weevilEurasian watermilfoil is an exotic aquatic plant that can take over lakes.  A native North American aquatic beetle, the milfoil weevil, eats Eurasian watermilfoil and sometimes suppresses watermilfoil infestations.  Unfortunately, the weevil does not contol watermilfoil in most lakes.  My research tested whether fish predation limits weevil populations.  I used a combination of field experiments and population monitoring to show that predatory sunfish can prevent weevil populations from becoming abundant enough to control watermilfoil.  So, by suppressing the herbivorous weevils, sunfish may facilitate watermilfoil infestations in lakes.

Relevant publications:

Ward, D.M. and R.M. Newman. 2006. Fish predation on Eurasian watermilfoil (Myriophyllum spicatum) herbivores and indirect effects on macrophytes. Canadian Journal of Fisheries and Aquatic Sciences 63: 1049-1057.